Hmmm, Pinnules look all folded up... or missing... Could you verify which? If they are folded it could just be irritated, if missing... Perhaps something has been nibbling at them and apparently for a while as the worm will just retract when bothered. I have not seen that before but it does look as though the pinnules are shriveled or retracted. But, the radiole are all out so that is good....
How long has it been like this? Have you done anything to your tanks that may correlate to what you have there?