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Ah, so you'd like to see what kind of neglect / abuse it takes to cause a Nitrifier community to lose its ability?While we are at it, I'd try to split them across three designs:
1. Churning
2. No churning
3. Churning, no oxygen
4. No churning, no oxygen (?)
(4) would be preferable, but I suppose not absolutely necessary to do. Not sure how you'd do anoxic conditions though.
The no-oxygen conditions could be achieved during the fish flake part of the experiment.
I think we have papers on subjecting nitrifying biofilms to these conditions, or are you hunting a particular hunch?